Tilly, a river otter, came to the Oregon Zoo in 2009 as an injured and orphaned pup. She birthed four pups while at the zoo.

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The Oregon Zoo said goodbye to a member of the otter family Wednesday. Tilly, the zoo’s oldest North American river otter, was humanely euthanized at age 16, due to advanced age, according to a press ...

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This is a very sad time, especially for Tilly’s care staff,” said Jen Osburn Eliot with the Oregon Zoo, in a statement.